And what about the X Y Z scale values? In the error message, LOD3 is referring to the High LoD model and Negative scale detected usually means that one or more of the X Y Z scale values in Blender > Object mode > Transforms side panel are set to a negative value. All three should be set to 1.000 before exporting    (and before using any modifiers or UV unwrapping in Blender). In the example below the Cube's X scale has been deliberately set to  -1.000  (negative 1.00) :
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